When Robert takes pity on a homeless family and takes them in, Emma’s horror is nothing compared to their shock when they discover the family have departed during the night, leaving behind their little daughter, Polly, as the adaptation of Flora Thompson’s magical memoir of her Oxfordshire childhood continues.

Lady Adelaide becomes aware of Polly’s existence and soon falls in love with her. Sir Timothy, however, is extremely wary of this. He knows how badly Adelaide wants a child, but for them to adopt Polly would be completely inappropriate. Sir Timothy takes Polly to the Post Office where an out-of-sorts Zillah is lifted by the bond she forges with the child. Adelaide realises she cannot keep Polly but follows Sir Timothy to the Post Office in order to say goodbye. As it becomes apparent that there’s no clear solution, little Polly’s future hangs in the balance.

Thomas, meanwhile, has been having trouble sleeping and his restlessness is keeping everyone else at the Post Office awake. When Laura and Dorcas realise that it is thoughts of Miss Ellison that are keeping him up, they decide to intervene by sending her a love poem on his behalf. Miss Ellison responds with a poem of her own. Her poem ends up in Sir Timothy’s possession, however, and events spiral out of control, making Dorcas wish she’d never intervened.

Back in Lark Rise, Queenie is increasingly concerned about Twister’s worsening delusions.

Robert Timmins is played by Brendan Cole, Emma Timmins by Claudie Blakley, Polly by Ruby Douglas, Lady Adelaide by Olivia Grant, Sir Timothy Midwinter by Ben Miles, Zillah by Liz Smith, Thomas Brown by Mark Heap, Laura Timmins by Olivia Hallinan and Dorcas Lane by Julia Sawalha.